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Zoology Publications from Victoria University of Wellington—Nos. 78, 79 and 80

Keys to Aid in the Identification of Marine Harpacticoid Copepods. Amendment Bulletin No. 5

page 1

Keys to Aid in the Identification of Marine Harpacticoid Copepods
Amendment Bulletin No. 5

Introduction

Included in this Bulletin is an important series of papers by Kunz on the Family Tetragonicipitidae that have caused me to completely recast the keys to this family. As in previous Bulletins (Wells, 1978, 1979, 1981, 1983) the page numbers in parentheses are those of the original Keys (Wells, 1976).

Key to Families

1.Couplet 9 (p. 6) can be misleading; amend by replacing "at outer distal corner" with "on outer border".
2.To make the key comprehensive for Family Tachidiidae
(a)add a new couplet to follow couplet 41 (p. 9) –
42a. Enp. P.1 of 2 segments Tachidiidae
Enp. P.1 of 3 segments 42b
(b)renumber couplet 42 as 42b,
(c)note that couplet 40 now leads to couplets 41 and 42a.

I am indebted to Morten Jødal and Michael Gee for suggesting these improvements.

Family Canuellidae

1.Add these new codons to KGG 1 (p. 18)
7:6:4:4/6:5:4:3/a/3/8 Canuellina tuba Por, 1983
5:4:4:4/6:5:4:4/a/3/7 Brianola vangoethemi Fiers, 1982
7:7:5:4/6:5:4:4/a/3/8 Scottolana glabra Fiers, 1982
S. dissimilis Fiers, 1982
S. uxoris Por, 1983
page 2
2. Scottolana antillensis Fiers, 1984c to codon for S. inopinata in KGG 1 (p. 18).
3.Fiers (1982) describes Canuella paenelantica n.sp. and states that he gives this name "because of the resemblance to Canuella elantica Por, 1967". Actually the correct name is elanitica (Por 1967: 106). Clearly paenelantica is an incorrect spelling due to a lapsus calami by Fiers and must be emended to paenelanitica (International Code of Zoological Nomenclature, 1985, Art. 32(c)(ii)).
4. Canuella paenelanitica Fiers, 1982 to codon for C. perplexa and C. furcigera in KGG 1 (p. 18).
5.Fiers (1982) erects Parasunaristes new genus. In KGG 1 (p. 19)
(a)Amend generic name of Sunaristes dardani, Ellucana curticaudata and E. chelicerata to Parasunaristes.
(b)Add P. cucullaris Fiers, 1982 to codon for P. dardani.
6.Fiers (1982) redescribes the female of Ellucana longicauda and (1984c) gives the first description of the male.

Family Cerviniidae

1. Eucanuella longirostrata Itô, 1983 to genus codon in KGG 1 (p. 21).
2. Cervinia plumosa Itô, 1983 to codon for C. tenuiseta and C. unisetosa in KGG 100 (p. 23).
3.Add these new codons to KGG 200 (p. 24)
5/2/5:7:7:7/4:5:6:5/>abd Cerviniopsis muranoi Itô, 1983
?/?/5:5:5:4/4:5:5:3/=abd C. minutiseta Itô, 1983

Family Ectinosomatidae

1. Arenosetella longiseta Kunz, 1983 to genus codon in KGG 1 (p. 28).
2. Pseudobradya beduina faialensis Kunz, 1983 to species codon in KGG 100 (p. 29).
3. Bradya (Bradya) pugiochaeta Arlt, 1983 to subgenus codon in KGG 200 (p. 31).
4.Hicks & Schriever (1983) erect the new genus Kliella to accommodate K. spinosa n.sp. and Halophytophilus ? triarticulatus Klie, 1949. As a consequence
(a)Add a new character-state to character 5 of KGG 1 (p. 27) — page 33:p = 3 segments; prehensile.
(b)The genus requires a new codon in KGG 1 (p. 28) — fa/3/bl:3/n/3:p.

Family Darcythompsoniidae

1.Mielke (1982) redescribes Darcythompsonia fairliensis (note that the legends for his Abb. 3 and 4 are transposed). As a consequence the following are amended codons for KGG 1 (p. 37)
1:1/1:2/4:4/0:0:0:1/3:4:4:4 D. scotti
1:1/0:1 or 1:2/4:4/0:0:0:1/3:4:4:4 D. fairliensis
1:1/?/3:7/0:0:0:1/3:4:4:4 D. inopinata
2. Leptocaris azoricus Kunz, 1983 requires a new codon in KGG 1 (p. 37) – 0:2/0:2/3:5/1:1:0:0/2-3:3:3:3.

Family Harpacticidae

Add Harpacticus longiantennata Apostolov & Petkovski, 1980 to footnote c of KGG 100 (p. 46).

Family Tisbidae

1. Tisbe spinulosa Bradford & Wells, 1983 and T. caymanensis Yeatman, 1984 to genus codon in KGG 100 (p. 49).
2.The new genus Volkmannia Boxshall, 1979, with its two new species, ***forficulata and attenuata, to codon for Tisbe and Bathyidia in KGG 100 (p. 49). Boxshall discusses these three genera and lists points of detail by which they can be separated (see also Wells, 1983).
3.KGG 300 (see Wells, 1983)
(a) Neotisbella gigas, a new genus and species by Boxshall (1979) requires a new codon — 8/4/7:8:8/5:6:5/2:2:2.
(b)As Apostolov & Petkovski (1980) reduce Zosime bathybia to a subspecies of Z. incrassata, delete Z. bathybia from footnote b.

Family Porcellidiidae

Porcellidium tapui Hicks & Webber, 1983 is added to this monogeneric family (see p. 12).

Family Clytemnestridae

Boxshall (1979) points out that the family name Pseudopeltidiidae is not valid and must be replaced by Clytemnestridae (see character 8 of page 4Key to Families (p. 6) and note 3 on p. 11).

Family Tegastidae

Syngastes spinifer Fiers, 1983 to genus codon in KGG 1 (p.56).

Family Thalestridae

1.KGG 1 (p. 58)
(a)Add these new codons
3:2/7:8:8/1:1/3/p:d Pseudotachidius horikoshii Itô, 1983
3:3/6:6:6/l:l/3/p:f P. minutus Itô, 1983
2:2/6:7:7/2:2/3/p:f Diarthrodes zavodniki Apostolov & Petkovski, 1980
(b) Pseudotachidius bipartitus pacificus Itô, 1983 to the species codon (see Wells, 1981).
2. Paradactylopodia striata Kunz, 1983 to genus codon in KGG 100 (p. 61). Kunz gives a key to this genus.

Family Parastenheliidae

Pallares (1982a) adds two new species, Parastenhelia costata and P. minuta to this monogeneric family (see p. 12).

Family Diosaccidae

1.The previously unknown male of Stenhelia (D.) minuta is described by Marinov & Apostolov (1981).
2.KGG 100 (p. 73)
(a)

amend column 3 of codon for Robert gurneya d to 0:0:0.

(b)delete codon for Robertgurneya smithi and add the species to footnote d.
(c)Add Haloschizopera bathyalis Schriever, 1984a to footnote c.
3.KGG 110 (p. 74)
(a) Amphiascoides breviarticulatus Kunz, 1983 requires a new codon - =Exp/se/7/5:5/3:5.
(b)Add Paramphiascella austroatlantica Pallares, 1982a to footnote b (p. 75).
4. Bulbamphiascus cibimae Pallares, 1982a to genus codon in KGG 230 (p. 80).
5.Kunz's (1983) specimens of Eoschizopera reducta show enhanced page 5setation; amend the species codon in KGG 300 (p. 85) to read — 0:1:1/3-4:3-4:2/4:4:5.
6.KGG 400 (p. 86)
(a)Add these new codons
4/6:8:8/8/d:?/lss S. (D.) intermedia Marinov & Apostolov, 1981
4/7:8:8/7/d:d/lss S. (D.) stephensoni Greenwood & Tucker, 1984
(b) S. (D.) islandica Schriever, 1982b to codon for S. (D.) cornuta and S. (D.) longipilosa (p. 86).
(c) Stenhelia (D.) adriatica Marinov & Apostolov, 1981 to codon for S. (D.) latisetosa (p. 87).
7.Add these new codons to KGG 700 (p. 91)
2/3/6:7:7/f:?/4:2 Pseudomesochra scheibeli Schriever, 1982b
2/3/4:4:4/f:?/4:6 Schizopera arconae Arlt, 1983.
8. Schizopera soyeri Kunz, 1983 to codon for S. arenicola and S. nichollsi in KGG 800 (p. 93).
9. Stenhelia (D.) noodti Schriever, 1982b requires a new codon in KGG 900 (p. 96) - 6:7:7/5:5:4/d.

Family Ameiridae

1.KGG 1 (p. 101)
(a)Add these new codons
3:2/3:3:3/3:3:2/5/0 Psyllocamptus minutus Pallares, 1982a
2:2/2:2:2/3:3:3/6/na Malacopsyllus hirsutus Itô, 1983
(b)Add Parapseudoleptomesochra helienica Pesce, 1981a and P. attirei Dumont, 1984 to footnote d on p. 103 (see Wells, 1978).
2.KGG 300 (p. 107)
(a)Add these new codons
s/0:0:0/1:1:1/3:4:4/2 Pseudameiropsis argentinus Pallares, 1982b
s/0:1:1/0:1:1/4:4:4/2 Nitocra baltica Arlt, 1983
s/0:0:0/1:1:1/5:4:5/1 N. mediterranea pontica Apostolov, 1980
s/0:0:0/l:l:l/3:3-4:4-5/1 N. lacustris azorica Kunz, 1983
(b)Note that on p. 108 the codon for N. mediterranea now refers to the nominate subspecies only.
(c)Note that in footnote b (p. 108) N. lacustris now refers to the subspecies lacustris, sinoi and the new subspecies pacificus Yeatman, 1983 only.
(d) Abyssameira reducta, a new genus and species by Itô (1983), page 6and Sarsameira knorri Reidenauer & Thistle, 1983 to codon for Sarsameira pendula.
3. Parameiropsis magnus Itô, 1983 to codon for P. peruanus in KGG 310 (p. 109).
4. Nitocra pseudospinipes Yeatman, 1983 requires a new codon in KGG 330 (p. 111) - 7:7:7/0:0:0/3-4:6/3:5-6.
5.KGG 400 (p. 114)
(a)Add these new codons
0:0:0/1:1:1/1:1:1/4:5:4/3:5/? Ameira faroerensis Schriever, 1982b
0:0:0/1:1:1/1:1:1/4:5:5/4:5/4:5 A. confluens Reddy, 1984
0:0:0/1:1:1/1:1:1/4:5:5/3-4:2/? Pseudameira antennulata Schriever, 1984a
0:0:0/1:1:1/1:1:1/3:3:3/4:2/? P. trisetosa Schriever, 1984a
(b)Kunz's (1983) specimens of Ameira tenella increase the degree of variation known for this species. As a consequence amend its codon (p. 115) to - 0:0:0/0:0-1:1/0:0.0/4:4-5:4/4:5-6/?
(c) Proameira thetiensis Pallares, 1982a to codon for P. simplex and P. arenicola (p. 116).
(d) Pseudameira mixta adriatica Apostolov & Petkovski, 1980 to species codon (see Wells, 1981).
6.Add these new codons to KGG 600 (p. 118)
4:4:5/1:1:1/3:2:2/3:4/? Nitocrella rhodiensis Pesce, 1983a
4:4:5/1:1:1/2:2:2/4:4/2:5 N. achaiae Pesce, 1981a
4:4:4/1:1:1/2:2:2/3:4/2:5 N. morettii Pesce, 1984
4:4:4/1:1:1/2:2:1/2-3:4/? N. juturna Cottarelli, 1975
4:4:4/1:1:1/1:2:1/2:4/? N. maggii Pesce, 1983b
7.KGG 700 (p. 120)
(a) Nitocrella skyrensis Pesce, 1981b requires a new codon — 4:4:4/1:1:1/2:2:2/2:4/2:4.
(b) Nitocrella somalica Dumont, 1981 to codon for N. petkovskii (see Wells, 1983).

Family Paramesochridae

1.KGG 1 (p. 124; see Wells, 1983)
(a)Add these new codons
3:3:3:3/3/3/5 Diarthrodella lancifera Kunz, 1983
2:3:3:3/3/3/6 D. galapagoensis Mielke, 1984b
2:2:3:3/3/2/7 D. chilensis Mielke, 1985
2:1:1:2/2/0/4 Scottopsyllus langi Mielke, 1984b
page 7
(b) Diarthrodella convexa Kunz, 1983 and D. neotropica Mielke, 1984b to codon for D. secunda.
(c) D. parorbiculata pacifica Mielke, 1984b to species codon.
(d) Rossopsyllus kerguelensis quellonensis Mielke, 1985 to species codon.
(d)Add Leptopsyllus punctatus and L. platyspinosus, new species by Mielke (1984b), to footnote e (p. 125).
(e)Kunz (1983) gives a key to Diarthrodella.
2.KGG 100 (p. 126)
(a) Paramesochra unaspina Mielke, 1984b requires a new codon — 4/4:4:2/0:0:0/2:3/0:3
(b) P. kunzi Mielke, 1984a to codon for P. dubia.
(c) P. helgolandica galapagoensis Mielke, 1984b to species codon.
3.KGG 200 (p. 127)
(a)Add these new codons
4:4:2/1:1:1/1:3/0:4 Kliopsyllus atlanticus Kunz, 1983
4:4:2/1:1:1/?/0:4 K. miguelensis Kunz, 1983
4:4:3/1:1:1/1:3/0:3 K. panamensis Mielke, 1984a
(b) Kliopsyllus constrictus pacificus Mielke, 1984a to species codon (p. 127).
(c)Add Kliopsyllus regulexstans and K. similis, new species by Mielke (1984b), to footnote b (p. 128).
(d)Add Kliopsyllus unguiseta Mielke, 1984b to footnote c (p. 128).
4.Mielke (1984a, b) describes Apodopsyllus panamensis, A. aberrans and A. arcuatus. These and other recent additions to this genus make interpretation of characters 3 and 4 of KGG 300 (p. 129) rather difficult. Replace KGG 300 with this new version.

KGG 300 - characters

1.P.1 Exp.
n = number of segments
2.P.1 Exp., distal (or only) segment
n = number of setae and/or spines
3.P.5♀
n = total number of setae and/or spines
4.P.5♂
n = total number of setae and/or spines
page 8

KGG 300

P.1 P.1 P.5♀ P.5♂
ExP. segs. ExP. distal seg. setae setae
2 4 6 4 Apodopsyllus vermiculiformis
A. panamensis
A. arcuatus
2 4 5 4 A. africanas listensis
2 4 4 4 A. africanas s.str.
A. schultzi
A. unguiformis
2 4 4 3 A. bermudensis
2 4 3(4?) 3 A. adaptatus
2 4 4 ? A. madrasensis
2 4 2 3 A. spinipes
2 4 ? 5 A. per plexus
1 5 5 5 A. camptus
1 5 4 4 A. littoralis
A. lynceorum
1 5 3-4 4 A. arenicolus
1 5 3 3 A. depressus

Family Tetragonicipitidae

Kunz (1984c) reviews the family and discusses its phylogeny. He places Fearia as a synonym of Tetragoniceps and declares Phyllopodopsyllus pirgos to be a juvenile male of P. thiebaudi.

Kunz (1984a, b) describes five new species of Phyllopodopsyllus (angolensis, petkovskii, geddesi, gertrudi, mielkei, with a subspecies m. californicus) and two new subspecies of P. longipalpatus — l. madagascarensis and l. hawaiiensis.

Pallares (1982a) redescribes Phyllopodopsyllus mossmani and places P. paramossmani in its synonymy.

Oniscopsis inabai Kitazima, 1983 is added to the genus.

These additions and amendments, together with the fact that the present key does not adequately deal with sexual dimorphism in Phyllopodopsyllus, have led me to construct the following new keys to the family to replace KGG 1 (pp. 131-133).

page 9

KGG 1 - characters

1.Cephalothorax ornamentation
p = process at posterior lateral corner present
a = process at posterior lateral corner absent
2.A.1, large dentiform projection
(Note: usually similar in both sexes, but may be reduced in the male and well developed in the female)
a = absent
1 = present on first segment
2 = present on second segment
3.P.1 Enp.
n = number of segments
4.P.2-P.4., distal (or only) segment
n:n:n = number of setae and spines on P.2, P.3 & P.4
5.P.2-P.4 Exp.3
n:n:n = number of setae and spines on P.2, P.3 & P.4

KGG 1

Cph. orn. A.1 proj. P.1 EnP. P.2-P.4 EnP. distal seg. setae P.2-P.4 Exp.3 setae
p 1 2 3:3:2 6:6:5 Laophontella typica
p 1 2 3:3:1 6:6:5 L. armata
p 1 2 3:3:1 6:6:4-5 L. horrida
a 1 2 4:4:4 6:7:7 Tetragoniceps prima
a 1 2 3:3:3 6:5:7 T. bergensis
a 1 2 3:3:3 5:5:5 T. bookhouti
a 1 2 3:3:3 5:4:7 T. truncata
a 1 2 3:3:3 4:4:5 T. malleolatus
a 1 2 3:3:? 6:5:4 T. longicaudata
a 1 2 ?:3:3 ?:5:6 T. scotti
a 1 2 2:3:3 5:5:5 T. bookhouti ♂
a 1 2 2:2:3 4:3:5 T. brownei ♀
a 1 2 2:?:3 5:?:6 T. arenicolous
a 1 2 ?:?:2 ?:?:7 T. dubius
a 1 2 ? ? T. brevicauda
a 2 3 3:3:2 5:4:5 Protogoniceps
a 2 2 2-3:1-3:1-3 4-5:4-6:4-7 KGG 100
a 2 2 ? ? Phyllopodopsyllus minor
a a 3 2:2:2 4:4:4 Pteropsyllus
a a 3 3:3:4 4:4:7 Diagoniceps trifidus
a a 3 3:3:3 5:5:8 D. menaiensis
a a 2 4:4:4-5 6:7:8 D. bocki
a a 2 4:4:4 6:7:8 D. kunzi page 10
a a 2 3:3:3 5:6:8 D. monodi
a a 2 2-3:2-3:2-4 4-5:4-6:5-7 KGG 200
a a 2 2-3:1:1 3:2:2 Oniscopsis
a a 2 2:2:3 4:3:5 Tetragoniceps brownei ♂
a a 2 ? ? Phyllo podo psyllus tristanensis

KGG 100-200 - characters

(The same five characters are used in both KGG)

1.P.2 Enp.2
n = number of setae
(Note: In the male the "setae" may include a long straight apophysis)
2.P.3 Enp.2
n:n = number of setae in ♀ and ♂
3.P.4 Enp.2
n:n = number of setae in ♀ and ♂
4.P.2-P.3 Exp.3
n:n = number of setae
5.P.4 Exp.3
n:n = number of setae in ♀ and ♂

KGG 100

P.2 Enp.2 P.3 Enp.2 ♀:♂ P.4 Enp.2 ♀:♂ P.2-P.3 Exp.3 P.4 Exp.3 ♀:♂
3 3:3 3:3 5:6 6:6 Phyllopodopsyllus chavei
3 3:3 3:3 4:4 5:5 P. medius
3 3:3 3:2 5:6 7:7 P. bermudae
3 3:3 3:2 5:6 7:6 P. setouchensis
P. mielkei s.str.
3 3:3 3:2 4:4 7:7 P. danielae
3 3:3 3:2 4:4 7:6 P. simplex
3 3:3 3:2 4:4 6:6 P. pauli
P. opisthoceratus
3 3:3 2:1 5:6 6:6 P. minutus
3 3:2 3:2 5:6 7:7 P. parafurciger
P. curtus
3 3:2 3:2 5:6 5-7:6 P. furciger
3 3:2 3:2 5:6 6:6 P. longicaudatus
3 3:2 3:2 4:4 6:6 P. borutzkyi
3 3:? 3:? 5:6 6:? P. mielkei californicus
3 3:? 3:? 4:4 6:? P. laticauda
3 3:? 2:? 4:4 6:? P. bahamensis
3 2:3 3:2 5:5 5:5 P. langi
3 2:2 2:2 5:6 7:7 P. parabradyi page 11
3(2♂) 3:2 3:2 5:6(5♂) 6:6 P. bradyi
2 3:3 3:2 4:4 6:6 P. paraborutzkyi
2 2:2 3:2 4:4 4:4 P. geddesi
2 2:2 2:2 4:4 5:5 P. hermani

KGG 200

P.2 Enp.2 P.3 Enp.2 ♀:♂ P.4 Enp.2 ♀:♂ P.2-P.3 Exp.3 P.4 Exp.3 ♀:♂
3 3:3 3:4 5:5 7:7 Diagoniceps laevis
3 3:3 3:3 4:4 7:6 Phyllopodopsyllus berrieri
3 3:3 3:2 5:6 7:6 P. briani
P. petkovskii
3 3:3 3:2 5:6(5♂) 7:6 P. angolensis
3 3:3 3:2 5:5 7:6 P. thiebaudi
3 3:3 3:2 5:5 7:5 P. gertrudi
3 3:3 3:2 4:4 7:6 P. punctatus
3 3:3 3:2 4:4 6-7:6 P. mossmani
3 3:? 3:? 5:6 6:? P. aegypticus
3 3:? 3:? 4:4 7:? P. hibernicus
P. laspalmensis
3(2♂) 3:3 3:2 4:4 6:6 P. hardingi
2 2:3 2:2 4:4 6:6 P. biarticulatus
2 2:? 3:? 4:4 6:? P. xenus
2 2:? 2:? 4:4 6:? P. paraxenus
P. longipalpatus sstr.
2(3♂) 2:2 2:2 4:4 7:5 P. l. madagascarensis
2(3♂) 2:2 2:2 4:4 7:6 P. l. hawaiiensis

Family Cylindropsyllidae

1.KGG 1 (p. 141)
(a) Arenopontia trisetosa Mielke, 1982 requires a new codon — s/2:na/f/1:1:2/p.
(b) Notopontia galapagoensis Mielke, 1982 to codon for Syrticola flandricus (see Wells, 1983).
2.KGG 200 (p. 143)
(a) Stenocaris baltica Arlt, 1983, known only from the male, requires a new codon — 2:2:2/ns/0:1/na.
(b)As S. pygmaea is a synonym of S. pontica delete the codon for the former and amend the codon for S. pontica to read 1-2:1:2/ns/1:1/6.
3.KGG 300 (p. 144)
(a) Leptastacus ctenatus and L. spatuliseta, new species by Mielke page 12(1982), require a new codon - p/2/0:0:1/3:4:5/1:1:0/1:1:2.
(b) Leptastacus dispinosus dispinosus Mielke, 1982 and L. d. panamensis Mielke, 1983a to codon for L. minutus.
4. Arenopontia peteraxi Mielke, 1982 requires a new codon in KGG 600 (p. 149) - 4/3:3:3/1-2:1-2:2/4:4/r.
5. Psammopsyllus stri and P. falciseta, new species by Mielke (1983b), require a new codon in KGG 700 (p. 151) - ♀/0/3:2:2/3:2:2/3:3.

Family Cletodidae

1.Schriever (1982a, 1984a) describes three species in a new genus, Thieliella, that he assigns to the Family Ancorabolidae. In a later note Schriever (in press) recognizes that this is not correct and that
(a) Thieliella endopodita must be transferred to Cletodes. It requires a new codon in KGG 600 (p. 167) — a/1:2/0.0/d:1:5/d:0:4.
(b) T. nordatlantica and T. reducta are synonyms of Monocletodes varians.
2.KGG 1 (p. 154)
(a)Schriever (1983, 1984b) describes five new species of Metahuntemannia which require these treatments
(i) M. pseudomagniceps: ♂ requires a new codon —
3:2/3:3:3/2:2/4:6:6:/4:3; ♀ to footnote b (p. 159).
(ii) M. bifida ♀ (male not known) requires a new codon —
3:1/3:3:3/0:0/4:4:4/na:na.
(iii) M. triarticulata requires new codons —
3:bs/3:3:3/2:2/5:6:6/l:2
3:bs/3:3:3/2:2/5:6:6/3:4
(iv) M. atlantica and M. arctica females (males are not known) to codon for M. gorbunovi ♀ and M. spinosa ♀ (p. 155).
(b)Add these new codons for new species by Schriever (1983)
3:1/3:3:3/1:1/7:7:7/5:5 Mesocletodes parabodini
3:2/3:3:3/2:2/6:6:5/3-4:4-5 M. variabilis
3:2/3:3:3/0:0/4:4:4/na:na Paranannopus phimosus
(c) Mesocletodes trisetosa Schriever, 1983 requires a new codon and a new footnote — 3:1/3:3:3/1:11/4:4:5/2:1
i)in this species P.4 Enp. is represented by a seta only.
(d) Heteropsyllus serratus Schriever, 1983 to codon for H. rostratus and H. masculus (p. 158).page 13
(e)Amend codon for Paranannopus elongatus (see Wells, 1981) to — 3:2/3:3:3/3:2/3:3:3/1:1. Note that this is also the codon for Cylindronannopus primus (p. 156).
(f)Schriever (1983) describes the previously unknown female of Paranannopus langi. As a consequence delete the male symbol from the species codon (p. 156).
3.After Arlt (1983)
(a)Amend columns 3 and 4 of codon for Enhydrosoma longifurcatum in KGG 500 (p. 165) to d-f:3:2-3/d-f:2:2.
(b)In KGG 600 amend column 4 of codon for Cletodes longicaudatus (p. 167) to d:2-3:5, and for Cletodes longifurca (p. 168) to d:2:5-6.
4.The genus name Echinocletodes Pallares, 1982a is preoccupied by Echinocletodes Lang, 1936 (Family Ancorabolidae). I propose its replacement by Rosacletodes n.gen., whose sole and type species is Echinocletodes kuehnemanni Pallares, 1982a. The species requires new codons in KGG 1 (p. 154) —
2:1/2:1:1/0:0/4:4:4/1:1
2:2/2:3:2/0:0/4:4:5/1:1

Family Laophontidae

1.

Cottarelli & Mura (1982) describe three new species of Afrolaophonte which require these treatments

(a) A. michaelae: ♂ to codon for A. pori in KGG 1-♂♂ (p. 188); ♀ requires a new codon in KGG 1-♀♀ (p. 172) —1:1:2/0:1:1/3:4/1/6.
(b) A. aequatorialis: ♂ to codon for A. schmidti (see Wells, 1983) in KGG 1-♂♂ (p. 188); ♀ requires a new codon in KGG 1-♀♀ (p. 172) - 1:1:1/0:1:1/4:5/1/6.
(c) A. leonis: ♂ unknown, ♀ to codon for A. schmidti in KGG 1-♀♀ (p. 172).

Note also that Cottarelli & Mura's legends for Figs. 3 and 4 are partially transposed.

2. Novolaophonte viatorum Cottarelli, Saporito & Puccetti, 1983, a new genus and species, requires new codons —
in KGG 1-♀♀ (p. 172) - 2:1s:1/0:0:0/4:3/1/5
in KGG 1-♂♂ (p. 188) - 3:3:3/1:1/1:3/1/3.
3. Paronychocamptus anomalus Reddy, 1984: ♀ to codon for Psammolaophonte spinicauda in KGG 1-♀♀ (p. 173), ♂ requires a new codon in KGG 400-♂♂ (p.194) - 6:6:5/1:1:1/4:4:3/0:0/p.page 14
4. Laophontina noodti Kunz, 1983 requires a new codon in KGG 1-♀♀ (p.172) — 3s:l:2/0:1:1/3:5/1/6. The male is unknown.
5.In KGG 600-♀♀ (p. 179) the codon for Paralaophonte innae (see Wells, 1983) should read — 6:7:7/1:1:1/0:0:0/4:5:4/0:0:0.
6.Fiers (1984a) removes Paralaophonte spinicauda to Laophonte and makes slight amendments to the description of the male. As a consequence
(a)amend genus name in KGG 1200-♀♀ (p. 182) and KGG 500-♂♂ (p. 195),
(b)add Laophonte spinicauda to footnote a of KGG 100-♂♂ (p.197).
7. Quinquelaophonte parasigmoides is placed incorrectly in KGG 1800-♀♀ (p.185); transfer this species and its codon to KGG 1400-♀♀ (p. 183). Similarly, transfer this species from footnote h to footnote i of KGG 1-♂♂ (p. 191).
8. Pseudonychocamptus marinovi Apostolov & Petkovski, 1980 requires a new codon in KGG 1900-♀♀ (p. 185) - 6:6:6/1:1:1/0:0:0/4:5:4/0:1:1. The male is unknown.

Family Latiremidae

Delamarella phyllosetosa Kunz, 1984d requires a new codon (with an accompanying note) in KGG 1 (see Wells, 1978) – 9/3:2/4/00/*

References

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page 16

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page 17

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page 18

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* the segments bear a number of short, broad structures, of which possibly two are true setae.